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A CPIE Notebook – Grasses and Sedges of Hawai‘i Page ii

Using a Dichotomous Key

If you are new to using or understanding a dichotomous key, the Introduction (and links provided) to Keys to the Aquatic Biota of the Hawaiian Islands will be useful. In particular, you should read the "Beginner's Orientation" section describing how our keys are set-up and how they can be used to arrive at a name for a biological specimen of interest. You may find a polychotomous key easier to use than a dichotomous key if you are just beginning to tackle grass/sedge indentifications. Try the identification keys at CPIE Multiaccess Key (MAK).

Specialized Terminology

Grasses and sedges present an additional challenge independent of your experience or familiarity in using a dichotomous key: botanists have developed a fairly specialized terminology for the structures of a grass and especially the grass flower (forget "petals" and "sepals"). The terminology applied to sedges is similar. You will need to gain an understanding of this terminology and A Key to the Common Grasses and Sedges of the Hawaiian Islands and Guam is set up to make this (hopefully) as painless as possible.

Information Sources / Corrections / Identification Help

This key is based upon the knowledge and personal experiences of the author, but also relies heavily on the references cited on Page x. All photographs and drawings are those of the author, unless credited otherwise. The keys are regularly updated—a significant advantage of a web-based presentation over a printed book. You can see when a particular page was last updated at the bottom of that page where the version number is given as a date. We very much welcome comments on the key and particularly concerning any couplet that is confusing or leads to an erroneous identification. You may send comments to the author (guinther@aecos.com). If you encounter a grass you cannot identify with this key you may place it in an envelope (there must be an inflorescence) with a note that includes location and island and your email address for a return answer (if I can identify it). Mail the specimen to "E. Guinther, 45-309 Akimala Pl. Kāne‘ohe, Hawai‘i, 76744".

All of the pages in this web document are regularly reviewed and additional species incorporated as time allows—consider the entire document comprises over 45 pages, many considerably longer than a normal book page. In the second half of 2023, the many grass species added to the Hawaiian flora by Faccenda (2022, 2023) have been slowly incorporated. In 2024, attention has been paid to improving the entire section on sedges, including expanding the integration of Guam sedges into the keys. A new feature is being added to the keys: in cases where it may be helpful to be aware of a similar appearing species described on a different page or is in a different genus, a link is provided in a "consider somewhat similar [link]..." statement placed below the species description.

Organization / Key Conventions

The organization of a dichotomous key can be either diagnostic (natural) or synoptical, the latter reflecting scientific classification based on evolutionary relationships, the former more appropriate to a key intended for less rigorous field or "home" use and of limited scope (to commonly occuring grasses and sedges within a specified area, as here limited to the Hawaiian Islands flora). A standard organizational technique in a diagnostic key is to sort out the most obvious structural differences early in the key and then progress through requiring ever finer details. This approach has been used on Page 1 of the key in sorting out woody bamboos (clearly distinct from other grasses) at couplet [10], exceptionally large grasses (like canes) at couplet [11], and then those grasses with other than clearly linear leaves at couplet [12]. What remains, beginning on Page 4, is the vast majority of grasses in Hawai‘i that require looking into their detailed structure in order to identify. These are grasss that might appear, to the untrained eye, pretty much alike.

The keys presented here differ from more typical published keys not just in using a less technical language but in providing descriptive detail on the final couplet for each species. This additional information (and accompanying figure(s) in some cases) allows the user to confirm arrival at a correct identification without having to seek a description of the species elsewhere. But we caution: the steps passed through in working towards a final identification are an important part of the species description not necessarily repeated in the final couplet. Working backwards several steps (using the couplet source link) can be used to review these characteristics.

A secondary goal of this guide is to move the user to an identification without requiring very close examination (using a magnifying lens, lupe, or microscope) of miniscule grass structures like glumes and ligules—"secondary" because, for many species of grasses and sdeges, confirmation of an identification is not possible without resorting to this level of anatomical detail. However, with experience, many of our common grasses and sedges can be recognized on sight. To this end, lists of the most common grasses and sedges in the Islands fitting a certain set of characteristics are given at appropriate places in the key. These short lists allow the user to "jump" to a likely conclusion, where a brief description and photograph(s) may provide sufficient confirmation of an identification. Another option would be to start at the multi-access key (MAK) to herbaceous plants and reduce the possible identifications to a manageable number. The possibles list of grasses or sedges resulting from using the MAK will have links to descriptive couplets and figures in this dichotomous key.

Although our keys follow the normally accepted layout for a dichotomous key, presentation as a webpage allows links to enable movement through the key. At each couplet, in parentheses next to the first choice, is a back link to the last couplet selected. In the far right column is a link to the next couplet based on the choice made. Using these links, the user can easily go backwards and forwards through their sequence of choices, and perhaps correct as necessary. Other conventions encountered are: 1) text following a curly brace "{" provides additional information that may be helpful but is not definitive of a species; 2) blue triangles () indicates that a link will leave the page (forwards or backwards).

Short-cuts to an Identification

Using the list of most likely grasses obtained from say the "multi-access key to grasses" is a short-cut of sorts for those not really into keying things out, but just hoping to ID a common grass plant. For those with a good knowledge of grasses, jumping ahead to a recognized genus or inflorescence type is also a short-cut. To facilitate this, the Table of Contents following will prove useful; but be cautious—skipping early steps in a key can easily mislead. Confirmation of a correct identification may require far more information than is provided in a terminal few couplets. If an attempt to identify a grass or sedge in hand has led to an unsatisfactory result, it is generally best to start over at the very first couplet. Clicking back through the links and more carefully considering the choices can result in a better outcome. Realize that any key will work only for those species considered in constructing the key. Consult the bottom of a page for species NOT considered (genus pages primarily).

TABLE OF CONTENTS
GRASSES, SEDGES, & RUSHES

The Grasses
     Basic characteristics of grasses
     Basic characteristics of grass flowers
     Start of Grass Key [couplet 10]
Page iii
Page iv
Page v
Page 1
The Sedges
     Basic characteristics of sedges (ToC)
     Basic characteristics of sedge flowers
     Start of Sedge Key [couplet 9]
Page vi
Page vi
Page viii
Page 30
The Rushes
     Start of Rush Key [couplet 30]
Page viii
Page 39

GRASSES
Very Large Grasses Page 2
Bamboos Page 3
Grass Inflorescence Types Page 4
Two or More Racemes or Spikes
    Genus Paspalum
    Genus Digitaria
Page 5
Page 6
Page 7
"Foxtail" Inflorescences
    Genus Cenchrus
Page 8
Page 9
Spike and Spike-like Inflorescences
    Genus Sporobolus
Page 10
Page 11
Digitate Inflorescences
    Genus Chloris
Page 12
Page 13
Paniculate Inflorescences
    Genus Dichanthium
    Genus Bothriochloa
    Genus Eragrostis
Page 14
Page 18
Page 19
Page 20
Lawn Grasses Page 21
Bibliography Page x
Index (List of Species) Page xi
Index (Common names) Page xii
CPIE Multi-Access Key (MAK)

Grasses Sedges


© 2012-24 AECOS, Inc. [ver. 240418] To next page Introduction / ToC ~ page ii
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